Parque Nacional Calilegua ( Ledesma, JUJUY, ARGENTINA )

Neita-Moreno, J.C. & B.C. Ratcliffe (2017).
The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile and Uruguay. Insecta Mundi, 0547: 1-36.